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Before the Events of Hellsing
At the beginning of the series, Seras' history is unknown, though hinted at through a very brief flashback in OVA I. Later in OVA IV, Walter tells Alucard that he had not met her family, as she had been orphaned. In Vol. 7 and OVA VII, Zorin Blitz casts an illusion on Seras, causing her to flashback to her very dark history. It is discovered that Seras' father, a police officer, "got in too deep" during an undercover mission. The group he had infiltrated traced him back to his home and brutally murdered him. Her mother, Mrs. Victoria, hid Seras in a closet and then confronted the criminals, who immediately killed her as well. A distraught Seras charged out and stabbed one of the criminals in the eye with a fork before being shot in the gut. The flashback ends with Seras seeing glimpses of the criminals raping her mother's corpse.
Sometime later, Seras was sent to live in an orphanage. The horrific murder of her parents caused her to react violently to bullies and other children. None of the teachers took kindly to her and discussed removing her from the orphanage if she continued to cause more trouble. In another flashback, Seras expresses her strongest wish - to become a police officer, despite being persuaded against doing so. It's unclear what happens to her afterward, whether she got adopted or not but she managed to become a police officer as she dreamed she would.
The Events of Hellsing
Seras was originally a police officer who, along with the rest of her squad, was sent into Cheddar Village to investigate a priest who was responsible for multiple homicides. A vampire disguised as (or possibly was formerly) a priest had taken control of the town and turned its inhabitants into Ghouls. The vampire took Seras hostage; unsure if she was a virgin or not, he planned to rape her to ensure she would turn into a ghoul after he drank her blood. When Alucard arrived, the priest used her as a human shield to coerce Alucard into letting him live. Alucard asked her if she was a virgin, and when she confirmed that she was, he shot through her to kill her captor. In her final moments of human life, she was given a choice: to die as a human or become a vampire. She chose to live and Alucard turned her into a vampire.
Given her condition, Seras was immediately employed by the Hellsing Organization after becoming a vampire. She was given a new uniform and almost immediately sent out on a mission by Integra with Alucard to eliminate some ghouls which had infested a household. While carrying out this mission she would come face to face with Father Alexander Anderson from the Vatican agency Iscariot who pierced her with numerous blessed blades from behind before proceeding to fight with Alucard.
During the course of this battle, Alucard's head was taken off by the priest and Seras, fearing her Master dead, took it and fled. As she pulled out the blades from her body Alucard's voice spoke to her, bidding her to drink his blood and become a true vampire. Seras, however, was too hesitant to do so before Anderson caught up to her, it was only the timely arrival of Integra and Alucard's subsequent resurrection which saved the situation.
Sometime later, Walter revealed to Seras that per Sir Integra's order, he disposed of her bed and replaced it with a coffin. It was then that Seras was also given her new weapon, the anti-tank cannon Hallconnen. The timing of this proved fortunate, as soon after the Hellsing Manor was attacked by two vampire brothers, Luke and Jan Valentine. The younger brother, Jan, led an army of ghouls that slaughtered the Hellsing troops and almost succeeded in reaching Integra and the visiting council members. Together Seras and Walter were able to stop this from happening. Walter led their attacks and Seras provided armed support, eventually managing to pin Jan down in a grapple she learned from her police training. After revealing the Hellsing troops were now Ghouls Jan broke free of her hold, sending Seras into the arms of the ghouls. She subsequently lost control and in a fit of bloodlust destroyed a number of them until Integra intervened.
Jan, now disabled by gunshot wounds at the hands of Integra and the council, died as an unknown entity set him alight from the inside, the vampire only giving a single clue as to who sent him, 'Millennium'. Meanwhile downstairs Alucard killed and consumed Luke Valentine.
Shortly afterwards, a confrontation with the Iscariot Section XIII leader Maxwell occurred at the British Imperial War Museum, during which Seras successfully managed to avert another fight between Alucard and Anderson by pretending to be a tour guide.
After the encounter, she has a run-in with The Wild Geese. Much later she is comically locked in her coffin by Alucard so she could be tricked into traveling in the jet to Brazil. When they arrive in Brazil, Seras awakens to Alucard telling her to prepare for battle with the Rio SWAT team.
Seras tries to protest but is berated by Alucard for her mercy toward mortals. During her master's fight with Tublucain Alahambra, she tries to shoot at him but fails. The two are then rescued by Pip Bernadotte after Alucard defeats Tublucain.
During a fight with Anderson, she tries to intervene but is scared into submission. At the meeting between Iscariot, Hellsing, and the Queen of England, she acts as a security guard of sorts.
During the Invasion of London, The Wild Geese and Seras are stationed at the Hellsing Organization headquarters while Alucard battles Rip Van Winkle and Integra encounters Alexander Anderson and the rest of Iscariot during the raid in London. When Zorin Blitz attacks the headquarters against the Major's wishes, Seras holds her off with the Hallconnen II. While Pip is impressed with her, he teasingly refers to her as "Girl," rather than calling her by her name (which Seras finds annoying). Moments before the battle in headquarters, Pip shares an intimate story with Seras about his time in London and all of the people he has met there. He tells her that he cannot forgive how they have "turn[ed] into corpses... eating [other] corpses," and encourages her to get revenge. This inspires Seras to take down the airship.
During the fight in headquarters, Seras (with Alucard's help) breaks Pip from the illusion cast by Zorin. However, the distraction allowed the Nazis to infiltrate the building. Seras takes the offense while Pip and the Wild Geese take the defense. Before going to their posts, Pip attempts (but fails) to steal a kiss from Seras who screams at him. Pip tells her "don't die," before she runs into battle, then later remarks on how she's a "good girl" and how "If you let a girl like her die, you're a disgrace as a man." Volume 7 opens up with a bloodbath, as Seras is devastated by how many of the Wild Geese have been violently butchered by the Nazis. She vows to beat all of them. Meanwhile, Pip and the others are being beaten back by the Nazis, though Pip is confident that Seras will come to their aid since she is "[that] kind of girl." Pip is wounded by the time Seras arrives but is overjoyed to see her. Before her fight with Zorin, he comments on how he wishes he could have kissed her.
Using her illusions, Zorin distracts Seras with the horrible memory of her father's death and the rape and murder of her mother. Zorin then slices off Seras' arm, stabs her in the back, and then blinds her with her scythe. Before Zorin can kill her, Pip hits her in the face with a rifle and shoots her. He throws Seras over his shoulders and attempts to escape. All the while, Seras begs him to leave her behind and save himself. He refuses but is fatally wounded when Zorin throws her scythe at his back. Blinded and weakened, Seras tries to find him when they fall to the ground, calling his name and feeling around for him. Pip waits until she's close enough before he moves in and kisses her. Seras is stunned by this but gives into his kiss. Pip laughs, happy that he finally stole a kiss from her, then tells her to feed on him. His final thoughts before dying were, "She's such a good girl. If I die protecting her, c'est la vie." Seras is devastated by his death, holding him in her arms while she screams in agony. It is only when Zorin insults Pip by referring to him as an insect that Seras is moved to rage and finally drinks Pip's blood. This restores her and allows her to kill the remaining vampires, including Zorin herself.
She then proceeds to save Integra from the Iscariot priests and meets up with Alucard on his return to London, astounded by his original form. Soon after she is present when Paladin Alexander Anderson dies and questions Walter for his betrayal of Hellsing. She and Integra are then invited to fight on the airship Deus Ex Machina along with The Major and The Captain. While Integra looks for the Major, Seras confronts the Captain. The two promptly engage in a swift gun battle, raining rounds at each other in a hectic fashion. As both are eventually disarmed, they move to hand-to-hand combat. The Captain now demonstrates how truly and utterly outmatched the young Vampire is, displaying his bestial wolf form, mind-staggering strength, use of mist-like intangibility, and blistering speed to absolutely dominate her in the initial exchange.
Viciously battering Seras, she is finally knocked through the floor, down into the storage room; a bullion horde of confiscated treasures from decades ago. Broken and hopeless, Seras is reinvigorated by her familiar, Bernadotte, and enters the fight again. After a quick flurry, The Captain coldly demonstrates the true extent of his mist form invulnerability, reflexive ability, and prowess to show Seras again how hopeless this battle is.
In an unexpected move, The Captain pointedly kicks an object into the young Vampires hand. Bemused, Seras notes that it is a silver tooth, ripped from the mouth of a Jewish captive many years earlier. Catching on immediately, Bernadotte calls the Captain a “good dog”, and notes that he deliberately led the fight here, and gave Seras the very tool of his undoing.
As the last werewolf gears up for a new assault, Seras realises this will be the final exchange, no matter what. The Captain launches at Seras and brutalises her once again. As they violently clash, Bernadotte urges and guides Seras, and soon they literally disarm each other, trading punches with enough force to messily explode entire limbs. Catching his swung kick, Seras pins the Captain’s leg, and Bernadotte plunges the silver tooth deep in his heart - delivering long overdue payback from some poor Jewish soul. Seras looks at the Captain and smiles widely after defeating him.
She then destroys the Major's shield, blowing away half his body in the process. Then she carries Integra from the zeppelin as it explodes.
Aftermath
30 years later, she declares Integra the winner of a fencing match and has seemingly developed a rivalry with Heinkel Wolfe. She then comes to her master's aid after hearing gunshots from her room but stops her attack when she finds Alucard back and alive after 30 years.
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